The three lions were related, and left orphaned when young cubs. They live in a wide open area within that fence, and are fed each day by Na'ankuse staff. So they know to appear at that spot in the fence every day. They were the first visit of our Carnivore Feeding Tour here at Na'ankuse, and already the shadows were very long, because the tour had been delayed to accommodate some guests arriving late. They never appeared, so the lighting was dim, but we did get a tour to ourselves - AND, we're going to get a whole make-up tour tomorrow.
One of the females was in heat and we saw the male mate twice with her. He's had a vasectomy because Na'ankuse has no license to breed lions, but he hasn't lost the desire.
After a brief few seconds, the female hisses, swats, and drives away the male.